Then: When Amanda asks the "Watermelon Guy" who Preston (or "Pre-stone") is, he and a friend offer her some not-so-helpful details: he's "sorta" tall, has hair, and wears t-shirts . . . sometimes.
Now: Segel has gone from supporting character to leading man, both on the big and small screen. He's lovably goofy as Marshall on How I Met Your Mother and showed off his comic chops in last year's The Five-Year Engagement.