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The story's protagonist, Surly squirrel (voiced by Will Arnett), is a controversial member of the city's animal community. Constantly embarking upon mischievous antics, he's excommunicated from his home, the park, after getting into trouble one too many times.
When the animals discover that there's a shortage of food and they won't be able to survive the upcoming Winter, two of the group's more honorable members, Grayson and Andie, are tasked with procuring nuts. They run into Surly on the streets and join forces for a heist that's full of constant twists and turns.