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It is revealed in the final few episodes that Eileen Hayward (Mary Jo Deschanel) had a long-ago affair with Benjamin Horne (Richard Beymer) and that he is actually Donna's (Lara Flynn Boyle) biological father, not Dr. Will Hayward (Warren Frost) as she had always believed. In the finale, Ben and Will come to blows in the Hayward living room, with the good doctor knocking Horne out cold.
Boyle is not appearing in the revival, but both Dr. Hayward and Ben Horne are, so perhaps we'll get to see how these two older gentlemen came to resolve their family drama — if they ever even did.