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While paging through a book at the Citadel, Gilly reads a certain maester's record of an annulment between Rhaegar Targaryen and his wife, and the new marriage that the maester officiated for Rhaegar afterwards. In other words, Rhaegar annulled his vows to Elia Martell and properly married Lyanna Stark before she gave birth to Jon, which means he's not a bastard at all. Jon is the rightful heir to the Iron Throne, even more so than Daenerys Targaryen.